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noun 
  1. (biochemistry) The material or substance on which an enzyme acts.
  2. (biology) A surface on which an organism grows or is attached e.g. the rock surface of a rockpool is the substrate for a sessile organism such as a limpet.
  3. An underlying layer; a substratum.
  4. (context, linguistics) An indigenous language that contributes features to the language of an invading people who impose their language on the indigenous population.
  5. (context, plating) A metal which is plated with another metal which has different physical properties.
  6. (context, construction) A surface to which a substance adheres.
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